What layers of the bowel wall comprise the valvulae conniventes

What layers of the bowel wall comprise the valvulae conniventes?

Valvulae conniventes (plicae circulares, folds of Kerckring) are composed of mucosa and submucosa. These folds increase the surface area of the small bowel. The villi of the small intestine are about 1 mm in cross-section and are just at the radiographic limits of resolution.

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